Mains water is reaching the DHW cylinder (at the top, i.e. the hot outlet), displacing the hot water within, and forcing it back up the cold feed (from the base of the cylinder) into the CSS.
For anyone who's still wondering why it does that, it's all down to pressure - the head of water in the vent is the same as that of in the cistern, so the water level will rise in both. It won't come out of the vent simply because it overflows from the cistern first.
My original comment was meant as a joke just in case you wondered.
However you still don't get it, The head of water in the vent is the same as that of the cystern (tank) That is until you backfeed it with MAINS pressure the water will go straight passed the cylinder (see drawing) and out the vent [/b]